Argyro Zenetos

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Argyro Zenetos

Research Director

Tel: +30-210-9856701
Fax: +30-210-9811713

Short CV

  • BSc in Natural Sciences, University of Athens (1971-75)
  • MSc in Paleoecology, Geology Dept, Univ. of St Andrews (1977-80, part time)
  • PhD in Marine Biology, Department of Biology, University of Athens (1981-84)

Dr. Argyro Zenetos is currently a research Director at the Institute of Biological Resources and Inland Waters HCMR, with a 29 year experience in the systematics and biodiversity of benthic macrofauna . Environmental impact studies and development of indicators related to pollution/disturbance from industrial effluent, (tannery, red mud, coarse metalliferous waste), oil spills, and trawling are included among her research activity. However, her main area of research is the Saronikos Gulf (which is being monitored for over 25 years) and the development of its benthic ecosystem. She is the co-ordinator of the Hellenic network on Aquatic Invasive Species (ELNAIS) and Chief Editor in the journal Mediterranean

Marine Science:

As a member of the SEBI2010 expert group on “trends in invasive alien species -see, She is responsible for marine alien species and has developed a Pan-European database which is updated to Dec 2013 under EEA contracts. Consultant to EEA and UNEP/MAP MEDPOL, UNEP/MAP on indicators for pollution under WFD.

Author of 110 peer-reviewed papers in scientific journals, 6 books and numerous participations in national and international conference, Zenetos is acting as a reviewer in many international journals while she is in the Advisory board of: Aquatic invasions (; Bioinvasions Records ( supervised 15 MSc and co-supervise(d)s 5 PhD theses and 1 post doc in progress.

Current activities include:


  • Simboura N, Zenetos A, Pancucci-Papadopoulou MA, Reizopoulou S, Streftaris N (2012) Hellenic Seas indicators for Sea-floor integrity under the European Marine Strategy Framework Directive: setting the thresholds and standards for Good Environmental Status. Mediterranean Marine Science 13:140-152
  • Zenetos Α, Gofas S, Morri C, Rosso A, Violanti D, et al. (2012) Alien species in the Mediterranean Sea by 2012.A contribution to the application of European Union’s Marine Strategy Framework Directive (MSFD). Part 2. Introduction trends and pathways. Mediterranean Marine Science 13:328-352
  • Katsanevakis S, Poursanidis D, Yokes B, Mačić V, Beqiraj S, Kashta L, Ramzi Sghaier Y, Zakhama-Sraieb R, Benamer I, Bitar G,  Bouzaza Z, Magni P, Bianchi CN, Zenetos A (2011) Twelve years after the first report of the crab Percnon gibbesi (H. Milne Edwards, 1853) in the Mediterranean: current distribution and invasion rates. Journal of Biological Research 16:224-236
  • Katsanevakis S, Stelzenmüller V, South A, Sørensen TK, Jones PJS, Kerr S, Badalamenti F, Anagnostou C, Breen P, Chust G, D’Anna G, Duijn M, Filatova T, Fiorentino F, Hulsman H, Johnson K,  Karageorgis AP, Kröncke I, Mirto S, Pipitone C, Portelli S, Qiu W, Reiss H, Sakellariou D, Salomidi M, van Hoof L, Vassilopoulou V, Vega Fernández T, Vöge S, Weber A, Zenetos A, ter Hofstede R (2011) Ecosystem-based marine spatial management: review of concepts, policies, tools, and critical issues. Ocean and Coastal Management 54:807-820
  • Katsanevakis S, Zenetos A, Mačić V, Beqiraj S, Poursanidis D,  Kashta L (2011) Invading the Adriatic: spatial patterns of marine alien species across the Ionian-Adriatic boundary. Aquatic Biology13:107-118
  • Tsiakkiros L, Zenetos A (2011) Further additions to the alien mollusc fauna along the Cypriot coast New opisthobranchia species. Acta Adriatica 52:115-124
  • Zenetos A, Gofas S, Verlaque M, Çinar ME, García Raso E, Azzurro E, Bilecenoğlu  M, Froglia C, Siokou I, Bianchi CN, Morri C, Sfriso A,  San Martin G, Giandgrande A, Katağan T, Ballesteros E,  Ramos-Esplá A, Mastrototaro F, Ocaña O,  Zingone A, Gambi MC, Streftaris N (2011) Errata to the Review Article (Medit. Mar. Sci. 11/2, 2010, 381-493): Alien species in the Mediterranean areas of the European Union’s  Marine Strategy Framework Directive (MSFD) by 2010. Part I.  Spatial distribution. Mediterranean Marine Science 12:509-514
  • Zenetos A, Katsanevakis S, Poursanidis D, Crocetta F, Damalas D, Apostolopoulos G, Gravili C, Vardala-Theodorou E, Malaquias M (2011) Additions and amendmentsto the marine alien biota of Greek waters (2010 update). Mediterranean Marine Science 12:95-120
  • Albayrak S, Balkis N, Balkis H, Zenetos A, Kurun A, Karhan SU, Caglar S, Balci M (2010) Golden Horn Estuary: Description of the ecosystem and an attempt to assess its ecological quality status using various classification metrics. Mediterranean Marine Science 11:295-313
  • Katsanevakis S, Poursanidis D, Issaris Y, Tsiamis K, Salomidi M, Maroulakis M, Kytinou E, Thessalou-Legaki M, Zenetos A (2010) The invasive crab Percnon gibbesi (H. Milne Edwards, 1853) [Crustacea: Decapoda: Plagusiidae] is spreading in the Aegean and Ionian Seas. Marine Biodiversity Records 3-e53:1-5
  • Raitsos D, Beaugrand G, Georgopoulos D, Zenetos A, Pancucci-Papadopoulou MA, Theocharis A, Papathanassiou E (2010) Global climate change amplifies the entry of tropical species into the Eastern Mediterranean Sea. Limnol. Oceanogr. 55:1478-1484
  • Zenetos A (2010) Trend in Aliens species in the Mediterranean. An answer to Galil, 2009 «Taking stock: inventory of alien species in the Mediterranean Sea». Biological Invasions 12:3379-3381
  • Zenetos A, Gofas S, Verlaque M, Çinar ME, Gracia Raso E, Azzurro E, Bilecenoglu M, Froglia C, Siokou I, Bianchi CN, Morri C, Sfriso A,  San Martin G, Giandgrande A, Katagan T, Ballesteros E,  Ramos-Espla A, Mastrototaro F, Ocana O,  Zingone A, Gambi MC, Streftaris N (2010) Alien species in the Mediterranean Sea by 2010. A contribution to the application of European Union’s Marine Strategy Framework Directive (MSFD). Part I. Spatial distribution. Mediterranean Marine Science 11: 381-493
  • Zenetos A, Ovalis P, Ceviker D (2010) On some Indo-Pacific boring endolithic Bivalvia species introduced in the Mediterranean with their host – Spread of Sphenia rueppelli A. Adams, 1850. Mediterranean Marine Science 11:201-207
  • Katsanevakis STsiamis K, Ioannou G, Michailidis N, Zenetos A (2009) Inventory of alien marine species of Cyprus (2009) Mediterranean Marine Science 10:109-133
  • Zenetos A, in press Trend in Aliens species in the Mediterranean. An answer to Galil, 2009 «Taking stock: inventory of alien species in the Mediterranean Sea», on line
  • Zenetos A, Konstantinou F, Konstantinou G (2009) Towards homogenization of the Levantine alien biota: Additions to the alien molluscan fauna along the Cypriot coast. Marine Biodiversity Records2:1-7
  • Zenetos A, Ovalis P, Kalogirou S (2009) Closing the gap:  Cerithium scabridum Philippi, 1848 found in the South Aegean (Greece, Mediterranean Sea). Journal of Biological Research-Thessaloniki 11:107-110
  • Zenetos A,Ovalis P, Vardala-Theodorou E (2009) The American piddock  Petricola pholadiformisLamarck, 1818 spreading in the Mediterranean Sea. Aquatic Invasions 4:385-387
  • Zenetos A, Pancucci-Papadopoulou MA,  Zogaris S, Papastergiadou E,Vardakas L, Aligizaki K, Economou AN (2009) Aquatic alien species in Greece: tracking sources, patterns and effects on the ecosystem. Journal of Biological Research-Thessaloniki 12:135-172
  • Kallianiotis A, Papaconstantinou C (2008) Artificial Reefs, 299-305p. In: State of Hellenic Fisheries. Ed. Papaconstantinou C, Zenetos A, Vassilopoulou V, Tserpes G, HCMR Publ., 466pp., Athens
  • Katsanevakis S, Lefkaditou E, Galinou-Mitsoudi S, Koutsoubas D, Zenetos A (2008) Molluscan species of minor commercial interest in Hellenic Seas: Distribution, exploitation and conservation status. Mediterranean Marine Science 9:77-118
  • Poursanidis D, Koutsoubas D, Issaris Y, Gerobasileiou V, Zenetos A (2008) New Records on the Opisthobranch fauna of the Greek Seas. JMBA Biodiversity Records 6374
  • Tsiamis K, Panayotidis P, Zenetos A (2008) Alien marine macrophytes in Greece : a review. Botanica Marina 51:237-246
  • Zenetos A, Meric E, Verlaque M,  Galli P,Boudouresque C-F, Giangrande A, ÇinarME, Bilecenoglu M (2008) Additions to the annotated list of marine alien biota in the Mediterranean with special emphasis on Foraminifera and Parasites. Mediterranean Marine Science 9:119-165
  • Zenetos A, Ovalis P, Houart R (2008) Eastward spread of Ergalatax junionae Houart, 2008 (Gastropoda, Mollusca) a recent alien in the Mediterranean. Journal of Biological Research 10:221-223
  • Daskos A, Zenetos A (2007) Additions to the knowledge of alien opisthobranchia of Greece. Aquatic Invasions 2:258-260
  • Louizidou P, Thessalou-Legaki M, Zenetos A (2007) First record of the Red Sea immigrant Chrysallida fischeri (Hornung & Mermod, 1925)  (Gastropoda: Pyramidellidae) from Greek watersAquatic Invasions 2:95-98
  • Zenetos A, Vassilopoulou V, Salomidi M, Poursanidis D (2007) Additions to the Marine Alien fauna of Greek Waters (2007 update). JMBA biodiversity records, 5928 on line
  • Albayrak S, Balkis H, Zenetos A, Kurun A, Kubanç C (2006) Ecological quality status of coastal benthic ecosystems in the Sea of Marmara. Marine Pollution Bulletin 52:790-799
  • Streftaris N, Zenetos A (2006) Alien Marine Species in the Mediterranean – the 100 ‘Worst Invasives’ and their Impact. Mediterranean Marine Science 7:87-118
  • Thessalou-Legaki M, Zenetos A, Kambouroglou V, Corsini-Foka M, Kouraklis P, Dounas C, Nikolaidou M (2006) The establishment of the invasive crab Percnon gibbesi (H. Milne Edwards, 1853) (Crustacea: Decapoda:Grapsidae) in Greek waters. Aquatic Invasions 1:133-136
  • Çevik C, Dogan A, Önen M, Zenetos A (2005) First record of the Indo-Pacific species Electroma vexillum (Reeve, 1857) (Mollusca; Bivalvia; Pterioidae) in the Eastern Mediterranean. JMBA2, biodiversity records, on line
  • Reizopoulou S, Zenetos A (2005) Summer resorts and benthic biodiversity in an open coastal area of the eastern Mediterranean. Mediterranean Marine Science 6:41-50
  • Simboura N, Zenetos A (2005) Increasing Polychaete biodiversity as a consequence of increasing research effort in Greek waters over a 70 years span with addition of new records and exotic species. Mediterranean Marine Science 6:75-88
  • Streftaris N, Zenetos A, Papathanassiou E (2005) Globalisation in marine ecosystems – The story of non indigenous marine species across European Seas. Oceanography & Marine Biology: An Annual Review 43:421-455
  • Zenetos A, Koutsoumbas D, Vardala-Theodorou E (2005) Origin and vectors of introduction of exotic molluscs in Greek Waters. Belgian Journal of Zoology 135:279-286
  • Zenetos A, Vardala-Theodorou E, Alexandrakis C (2005) Update of the Marine Bivalvia Mollusca check list in Greek Waters. Journal of the Marine Biological Association UK 85:993-998
  • Ståhl H, Hall POJ, Tengberg A, Josefson AB, Streftaris N, Zenetos A, Karageorgis AP (2004) Respiration and sequestering of organic carbon in shelf sediments of the oligotrophic northern Aegean Sea. MEPS 269:33-48
  • Zenetos A, Chadjianestis I, Lantzouni M, Simboura M, Sklivagou E, Arvanitakis G (2004) The Eurobulker oil spill: mid-term changes of some ecosystem indicators. Marine Pollution Bulletin 48:122-131
  • Gofas S, Zenetos A (2003) Exotic molluscs in the Mediterranean basin: current status and perspectives. Oceanography & Marine Biology: an annual review 41:237-277
  • Bogdanos C, Simboura N, Zenetos A (2002) The benthic fauna of Geras Gulf, (Lesvos isl. Greece): inventory, distribution and some zoogeographical considerations. Hellenic Zoological Archives 6:1–22
  • Simboura N, Zenetos A (2002) Benthic indicators to use in ecological quality classification of Mediterranean soft bottom marine ecosystems, including a new Biotic index. Mediterranean Marine Science 3:77-111
  • Zaitsev YuP, Alexandrov BG, Berlinsky NA, Zenetos A (2002) The Black Sea an oxygen-poor sea. In: Europe’s biodiversity – biogeographical regions and seas. Environmental issue report  Published by  EEA (European Environment Agency) Copenhagen 2002
  • Zenetos A, Siokou-Frangou I, Gotsis-Skretas O, Groom S. (2002) The Mediterranean Sea – blue oxygen-rich, nutrient-poor waters. In: Europe’s biodiversity – biogeographical regions and seas. Environmental issue report  Published by  EEA (European Environment Agency) Copenhagen 2002
  • Zenetos A, Simboura N (2001) Soft bottom benthic indicators. Rapp. Comm. Int. Mer Medit. 36:339
  • Zenetos A, Revkov NK, Konsulova Ts, Sergeeva N, Simboura N, Todorova VR, Zaika VE (2000) Coastal Benthic Diversity In The Black And Aegean Seas. Mediterranean Marine Science 1-2:105-117
  • Kallonas M, Zenetos A, Gofas S (1999) Notes on the ecology and distribution of microbivalvia in Greek waters. Conchiglia 291:11-20
  • Murina GVV, Pancucci–Papadopoulou MA, Zenetos A (1999) The phylum Sipuncula in the eastern Mediterranean: composition, ecology, zoogeography. Journal of the Marine Biological Association of the UK 79:821-830
  • Pancucci-Papadopoulou MA, Simboura N, Zenetos A, Thessalou-Legaki M, Nicolaidou A (1999) Benthic invertebrate communities of NW Rodos island (SE Aegean Sea) as related to hydrological and geographical location. Israel Journal of Zoology 45:371-393
  • Simboura N, Zenetos A, Pancucci-Papadopoulou MA, Thessalou-Legaki M, Papaspyrou S (1998) A baseline study on benthic species distribution in two neighbouring gulfs, with and without access to bottom trawling. Marine Ecology 19:293-309
  • Stergiou C, Christou N, Georgopoulos D, Zenetos A, Souvermetzoglou K (1997). Hellenic Seas. Oceanography & Marine Biology: an annual review 35:415-538
  • Zenetos A (1997) Diversity of marine Bivalvia in Greek waters: Effects of geography and environment. Journal of the Marine Biological Association of the United Kingdom 77:463-472
  • Zenetos A, Christianidis S, Pancucci MA, Simboura N, Tziavos Ch (1997) Oceanologic study of an open coastal area in the Ionian Sea with emphasis on its benthic fauna and some zoogeographical remarks. Oceanologica Acta 20:437-451
  • Zenetos A (1996) Classification and interpretation of the established Mediterranean biocoenoses based solely on bivalve molluscs. Journal of the Marine Biological Association of the United Kingdom 76:403-416
  • Simboura N, Zenetos A, Panayotidis P, Makra A (1995) Changes of benthic community structure along an environmental pollution gradient. Marine Pollution Bulletin 30:470-474
  • Simboura N, Zenetos A, Thessalou-Legaki M, Pancucci MA, Nicolaidou A (1995) Benthic communities of the infralittoral in the N. Sporades (Aegean sea): a variety of biotopes encountered and analysed. P.S.Z.N.I. Marine Ecology 16:283-306
  • Zenetos A (1994) First finding of Scapharca demiri (Piani, 1981) in the North Aegean Sea. La Conchiglia (Rome) 271:37-38
  • Zenetos A (1994) Records of bivalvia in Greek Waters. Biologia Gallo-hellenica 22:293-304
  • Nicolaidou A, Zenetos A, Pancucci M-A, Simboura N (1993) Comparing ecological effects of two different types of pollution using multivariate techniques. Mar. Ecol. 14:113-128
  • Pancucci MA, Panayotidis P, Zenetos A (1993) Morphological changes in sea-urchin populations as a response to environmental stress. In: Quantified Phenotypic Responses in Morphology and Physiology (Aldrich, J.C. ed.), JAPAGA, Ashford : 247-257
  • Papathanassiou E, Zenetos A (1993) A case of recovery in benthic communities following a reduction in chemical pollution in a Mediterranean ecosystem. Marine Environmental Research 36:131-152
  • Zenetos A (1992) Οn the bivalvia of Crete: a reply to Barash and Zenziper. La Conchiglia 263:21-22
  • Zenetos A (1991) Re-evaluation of numerical methods for delimiting biofacies and biotopes in an estuarine  environment. Lethaia 24:13-26
  • Zenetos A, Bei F, Nicolaidou A (1991) Erratic occurrence of benthic fauna in a shallow Mediterranean area: An  indirect effect of manmade disturbance. Marine Pollution Bulletin 22:618-662
  • Zenetos A, Papathanassiou E, Van Aartsen JJ (1991) Analysis of benthic communities in the Cyclades plateau (Aegean Sea) using ecological and paleoecological data sets. Marine Ecology-Pubblicazioni Della Stazione Zoologica Di Napoli I 12:123-137
  • Nicolaidou, A., F. Bourgoutzani and A. Zenetos, 1990. Effects  of pollution on the distribution of molluscs and polychaetes  in Messologhi lagoon and adjacent areas.  Map Technical Reports Series No. 40: 45-61, UNEP, Athens.
  • Pancucci MA, Zenetos A (1990) On the Echinoderm-fauna of the Geras Gulf. Acta Adriatica 31:293-300
  • Zenetos A (1990) Discrimination of autochthonous  versus  allochthonous assemblages in the Eden estuary, Fife,  Scotland. Estuarine, Coastal and Shelf Science 30:525-540
  • Zenetos A, Panayotidis P, Simboura N (1990) Etude des peuplements benthiques de substat meuble au large du debouche en  Mer du Grand Collecteur d’ Athenes. Revue International Oeanographie Medical 97-98:57-71
  • Nicolaidou A, Pancucci MA, Zenetos A (1989) The Impact of Dumping Coarse Metalliferous Waste on the Benthos in Evoikos Gulf, Greece. Mar.Pollut Bull. 20: 28-33
  • Pancucci MA, Zenetos A (1989) Infralittoral macrobenthos of the Patras Gulf and Ionian Sea: Echinodermata. Cahiers de Biologie Marine 30:217-226
  • Zenetos A, Papathanassiou E (1989) Community Parameters  and  Multivariate Analysis as a Means of Assessing the  Effects of Tannery Effluents on the Macrobenthos. Mar. Pollut. Bull. 20:176-181
  • Friligos N, Zenetos A (1988) Elefsis Bay Anoxia: Nutrient  Conditions and Benthic Community Structure. Marine Ecology-Pubblicazioni Della Stazione Zoologica Di Napoli I 9:273-290
  • Nicolaidou A, Bourgoutzani F, Zenetos A, Guelorget O, Perthuisot J-P (1988) Distribution of Molluscs and Polychaetes in coastal lagoons in Greece. Estuar. coast. Shelf Sci. 26:337-350
  • Zenetos A, Bei F (1988) Preliminary studies on the community structure of the macrozoobenthos in Atalanti Bay, Greece. Biologia Gallo-hellenica 13:21-24
  • Van Aartsen JJ, Zenetos A (1987) The genus  Clathromangelia Monterosato, 1884, in the Mediterranean, with the  description of Clathromangelia ferhi n.sp.  La  Conchiglia (Rome) 222-223:10-11
  • Zenetos A, Bogdanos C (1987) Benthic community structure as a tool in evaluating effects of pollution in Elefsis Bay. Thalassographica 10:7-21
  • Pancucci MA, Zenetos A (1986). Gli Echinodermi del golfo  di Patrasso. Biologia Gallo-hellenica12:189-195
  • Zenetos A, Bogdanos C (1986) Assessment of degree of pollution on the basis of macrozoobenthos in Elefsis bay. Rapp. Comm. Int. Mer Medit. 30:22
  • Zenetos A, Bratits C (1986) On the Prosobranchia (Mollusca, Gasteropoda) of the infralittoral zone along the Patras Gulf. Biologia Gallo-hellenica 12:167-172
  • Castritsi-Catharios J, Ganias G, Panagopoulos D, Zenetos A (1985) Bryozoaires cotiers du Golfe de Patras septentrional. Rapp Comm int Mer Medit 29:347-348
  • Pancucci MA, Zenetos A (1985) Contribution to the knowledge of echinoderms in the eastern Ionian sea. Rapp. Comm. Int. Mer Medit. 29:295-296
  • Thessalou-Legakis M, Zenetos A (1985) Autoecological Studies on the Thalassinidae (Crustacea, Decapoda) of the Patras Gulf and Ionian Sea (Greece). Rapp. Comm. Int. Mer Medit. 29:309-312
  • Thompson TE, Jarman GM, Zenetos A (1985) Infralittoral macrobenthos of the Patras Gulf and Ionian Sea: Opisthobranch Molluscs.  Journal of Conchology 32:71-95
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